The geology and biology of Grand Canyon are both vast in scope and deeply intertwined, but while the story of its rock is writ large, much of its highly diverse life is cryptic or, in the case of dragonflies, fast-moving and difficult to see. Therefore, it requires considerable effort to observe and understand life's particularity and spatial distribution in relation to the landscape. Over the past century, research by a committed group of scientists and naturalists has revealed much about the Canyon's plant life, fish, and birds, and has provided moderately good documentation of its other vertebrates. However, we have only just begun to plumb the depths of how the world's most famous large, deep canyon influences the myriad of invertebrate species that scuttle, blow, crawl, and zoom through its intimate crevices and gaping chasms, and how life persists here. Spectacular advances in scientific information over the past several decades provide ever-improving insights into relationships among the many species that live around us. Nonetheless, the literature on invertebrate biology remains primarily the domain of specialists and has not been made widely available to the public. Therefore, exploration, revelation, and outreach have been important activities for us to communicate to the public the richness of our natural biological heritage. Our aim here is to better understand and care for the beauty, complexity, and diversity of the varied life forms existing around us in this marvelous landscape and everywhere, and to provide a baseline for Grand Canyon against which to evaluate future biological changes. We hope this book conveys some of our appreciation of that beauty and sense of wonder.